REspectfully, with intent to educate not prove anyone wrong. Laquer is an interesting material. when you spray a coat of laquer, a large portion of the solvent does not evaporate immediatlyl, it actually makes its way from the new coat down into the previous coats all the way to the substrate. when it reachs that, then it reverses direction and evaporates out. Or it continues through the substrate in this case balsa and can evacuate the other side I imagine. I know that on cars, sheetmetal and filler, laquer will dry much faster over sheetmetal than over layers of built up primer because the solvent is absorbed less by the underlying layers. NOw that being said, if this is in fact enamel primer, its apparantly staying tight enough to survive partially. However over the fillets what could be happening is that there is no layers of material under it to absorb the solvent so itis bubbling the surface. where there are other layers of laquer under the primer, it wil absorb a portion of the solvent allowing a slower evaporation and less likely hood for bubblling to occur. The physical reason that material bubbles is because it has dryed and established its "finished dimension" IOW it has decided how much space it will occupy so to speak. then when you introduce solvent as in laquer into the dryed volume of enamel, it does not reflow like laquer does, it cannot vary its volume to accomodate the influx of solvent and in trying to it swells underneath causign surface tension to get very high causing the paint layer to flex and swell, poof bubbles.
Hope this explains a bit.
of course this is all theoreticall because we arent there watching it happen but it sure sounds like laquer over enamel.