For a uniflow tank, the point where the uniflow opens into the tank. That tank doesn't look like a uniflow, however, so for that one I'd say the centerline of the tank.
For profiles you often want the tank about 1/8" higher than the hole in the spraybar to start, and then adjust from there. Without muffler pressure, however, I think it's all moot: check back with us when you give up and go to a uniflow.